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Edexcel GCSE Biology Past Papers
Welcome to our page featuring a collection of Edexcel GCSE Biology Past Papers. These papers are essential resources for students preparing for their GCSE Biology exams. Practicing with these past papers can help improve your exam technique, enhance your knowledge and understanding of the subject matter, and boost your confidence. Our page provides access to a range of past papers, including those from the official Edexcel website, and offers tips and advice on how to use them effectively. Start practicing early to increase your chances of exam success.
1. Key Concepts in Biology
- 1 | Cell Basics Question Paper Mark Scheme
- 1 | Enzymes Question Paper Mark Scheme
- 1 | Cell Transport (2018 - 2021) Question Paper Mark Scheme
- 1 | Cells and Microscopy (2018 - 2021) Question Paper Mark Scheme
- 1 | Enzymes (2018 - 2021) Question Paper Mark Scheme
2. Cells and Control
- 2 | Brain and the Eye Question Paper Mark Scheme
- 2 | Cell Division Question Paper Mark Scheme
- 2 | Cell Division and Growth (2018 - 2021) Question Paper Mark Scheme
- 2 | Specialised Cells & the Nervous System Question Paper Mark Scheme
- 2 | The Eye (2018 - 2021) Question Paper Mark Scheme
- 2 | The Nervous System (2018 - 2021) Question Paper Mark Scheme
3. Genetics
- 3 | DNA & Protein Synthesis Question Paper Mark Scheme
- 3 | Inheritance Question Paper Mark Scheme
- 3 | Sexual & Asexual Reproduction Question Paper Mark Scheme
- 3 | Inheritance (2018 - 2021) Question Paper Mark Scheme
- 3 | Reproduction (2018 - 2021) Question Paper Mark Scheme
- 3 | The Genome (2018 - 2021) Question Paper Mark Scheme
- 3 | Variation and Human Genome (2018 - 2021) Question Paper Mark Scheme
4. Natural Selection and Genetic Modification
- 4 | Genetic Engineering & Selective Breeding Question Paper Mark Scheme
- 4 | Natural Selection & Evolution Question Paper Mark Scheme
- 4 | Classification(2018 - 2021) Question Paper Mark Scheme
- 4 | Evolution(2018 - 2021) Question Paper Mark Scheme
- 4 | Genetic Engineering(2018 - 2021) Question Paper Mark Scheme
5. Health, Disease and the Development of Medicines
- 5 | Human Disease Question Paper Mark Scheme
- 5 | Human Disease(2018 - 2021) Question Paper Mark Scheme
- 5 | Immune Systems Question Paper Mark Scheme
- 5 | Immune Systems(2018 - 2021) Question Paper Mark Scheme
6. Plant Structures and their Functions
- 6 | Leaf Structure & Function Question Paper Mark Scheme
- 6 | Leaf Structure & Adaptations(2018 - 2021) Question Paper Mark Scheme
- 6 | Photosynthesis & Plant Growth Question Paper Mark Scheme
- 6 | Photosynthesis (2018 - 2021) Question Paper Mark Scheme
- 6 | Plant Hormones Question Paper Mark Scheme
- 6 | Plant Hormones (2018 - 2021) Question Paper Mark Scheme
- 6 | Transport in Plants Question Paper Mark Scheme
- 6 | Transport in Plants (2018 - 2021) Question Paper Mark Scheme
7. Animal Coordination, Control and Homeostasis
- 7 | Homeostasis Question Paper Mark Scheme
- 7 | Homeostasis(2018 - 2021) Question Paper Mark Scheme
- 7 | Hormonal Control(2018 - 2021) Question Paper Mark Scheme
- 7 | Menstrual Cycle & Pregnancy Question Paper Mark Scheme
- 7 | Menstrual Cycle and Fertility(2018 - 2021) Question Paper Mark Scheme
- 7 | Urinary Systems Question Paper Mark Scheme
8. Exchange and Transport in Animals
- 8 | Circulatory System Question Paper Mark Scheme
- 8 | Respiration(2018 - 2021) Question Paper Mark Scheme
- 8 | Respiration Question Paper Mark Scheme
- 8 | The Circulatory System(2018 - 2021) Question Paper Mark Scheme
9. Ecosystems and Material Cycles
- 9 | Cycles within Ecosystems Question Paper Mark Scheme
- 9 | Cycles within Ecosystems(2018 - 2021) Question Paper Mark Scheme
- 9 | Feeding Relationships & Organisms in the Environment Question Paper Mark Scheme
- 9 | Humans & the Environment Question Paper Mark Scheme
- 9 | Humans & the Environment(2018 - 2021) Question Paper Mark Scheme
- 9 | Trophic Levels & Ecosytems(2018 - 2021) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2019 (New Spec)
- 2019 | QP - Paper 1 (F) Question Paper Mark Scheme
- 2019 | QP – Paper 1 (H) Question Paper Mark Scheme
- 2019 | QP – Paper 2 (F) Question Paper Mark Scheme
- 2019 | QP – Paper 2 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers May 2019 (New Spec)
- 2019 | QP – Paper 1(F) Question Paper Mark Scheme
- 2019 | QP – Paper 1(H) Question Paper Mark Scheme
- 2019 | QP – Paper 2(F) Question Paper Mark Scheme
- 2019 | QP – Paper 2(H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2018 (New Spec)
- 2018 | QP – Paper 1(F) Question Paper Mark Scheme
- 2018 | QP – Paper 1(H) Question Paper Mark Scheme
- 2018 | QP – Paper 2(F) Question Paper Mark Scheme
- 2018 | QP – Paper 2(H) Question Paper Mark Scheme
Edexcel GCSE Biology Specimen Past Papers (New Spec)
- 1 | QP - Paper 1(F) Question Paper Mark Scheme
- 2 | QP - Paper 1(H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers January 2018 (Old Spec)
- 2018 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2018 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2018 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2018 | QP - Unit B2 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2017 (Old Spec)
- 2017 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2017 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2017 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2017 | QP - Unit B2 (H) Question Paper Mark Scheme
- 2017 | QP - Unit B3 (F) Question Paper Mark Scheme
- 2017 | QP - Unit B3 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2016 (Old Spec)
- 2016 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2016 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2016 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2016 | QP - Unit B2 (H) Question Paper Mark Scheme
- 2016 | QP - Unit B3 (F) Question Paper Mark Scheme
- 2016 | QP - Unit B3 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2015 (Old Spec)
- 2015 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2015 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2015 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2015 | QP - Unit B2 (H) Question Paper Mark Scheme
- 2015 | QP - Unit B3 (F) Question Paper Mark Scheme
- 2015 | QP - Unit B3 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2014 (Old Spec)
- 2014 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2014 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2014 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2014 | QP - Unit B2 (H) Question Paper Mark Scheme
- 2014 | QP - Unit B3 (F) Question Paper Mark Scheme
- 2014 | QP - Unit B3 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2013 (Old Spec)
- 2013 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2013 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2013 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2013 | QP - Unit B2 (H) Question Paper Mark Scheme
- 2013 | QP - Unit B3 (F) Question Paper Mark Scheme
- 2013 | QP - Unit B3 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers March 2013 (Old Spec)
Edexcel GCSE Biology Past Papers November 2012 (Old Spec)
- 2012 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2012 | QP - Unit B1 (H) Question Paper Mark Scheme
- 2012 | QP - Unit B2 (F) Question Paper Mark Scheme
- 2012 | QP - Unit B2 (H) Question Paper Mark Scheme
Edexcel GCSE Biology Past Papers June 2012 (Old Spec)
Edexcel gcse biology past papers march 2012 (old spec), edexcel gcse biology past papers november 2011 (old spec).
- 2011 | QP - Unit B1 (F) Question Paper Mark Scheme
- 2011 | QP - Unit B1 (H) Question Paper Mark Scheme
Free GCSE Biology Tutorials
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1-to-1 lessons tailored to Edexcel GCSE Biology Past Papers
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Edexcel Biology Past Papers
Pearson Edexcel GCSE Biology past exam papers. If you are not sure what tier you are sitting foundation or higher check with your teacher. You can download the papers and marking schemes by clicking on the links below.
June 2022 Pearson Edexcel Biology Past Exam Papers (1BI0)
June 2022 Biology Paper 1 (1BI0/1F) – Foundation Tier Download Paper – Download Marking Scheme
June 2022 Biology Paper 1 (1BI0/1H) – Higher Tier Download Paper – Download Marking Scheme
June 2022 Biology Paper 2 (1BI0/2F) – Foundation Tier Download Paper – Download Marking Scheme
June 2022 Biology Paper 2 (1BI0/2H) – Higher Tier Download Paper – Download Marking Scheme
November 2021 Pearson Edexcel Biology Past Exam Papers (1BI0)
November 2021 Biology Paper 1 (1BI0/1F) – Foundation Tier Download Paper – Download Marking Scheme
November 2021 Biology Paper 1 (1BI0/1H) – Higher Tier Download Paper – Download Marking Scheme
November 2021 Biology Paper 2 (1BI0/2F) – Foundation Tier Download Paper – Download Marking Scheme
November 2021 Biology Paper 2 (1BI0/2H) – Higher Tier Download Paper – Download Marking Scheme
November 2020 Pearson Edexcel Biology Past Exam Papers (1BI0)
November 2020 Biology Paper 1 (1BI0/1F) – Foundation Tier Download Paper – Download Marking Scheme
November 2020 Biology Paper 1 (1BI0/1H) – Higher Tier Download Paper – Download Marking Scheme
November 2020 Biology Paper 2 (1BI0/2F) – Foundation Tier Download Paper – Download Marking Scheme
November 2020 Biology Paper 2 (1BI0/2H) – Higher Tier Download Paper – Download Marking Scheme
Some of the above papers are labelled June 2020
June 2019 Pearson Edexcel Biology Past Exam Papers (1BI0)
June 2019 Biology Paper 1 (1BI0/1F) – Foundation Tier Download Paper – Download Marking Scheme
June 2019 Biology Paper 1 (1BI0/1H) – Higher Tier Download Paper – Download Marking Scheme
June 2019 Biology Paper 2 (1BI0/2F) – Foundation Tier Download Paper – Download Marking Scheme
June 2019 Biology Paper 2 (1BI0/2H) – Higher Tier Download Paper – Download Marking Scheme
June 2018 Edexcel Biology Past Exam Papers (1BI0)
June 2018 Biology Paper 1 (1BI0/1F) – Foundation Tier Download Paper – Download Marking Scheme
June 2018 Biology Paper 1 (1BI0/1H) – Higher Tier Download Paper – Download Marking Scheme
June 2018 Biology Paper 2 (1BI0/2F) – Foundation Tier Download Paper – Download Marking Scheme
June 2018 Biology Paper 2 (1BI0/2H) – Higher Tier Download Paper – Download Marking Scheme
June 2017 Edexcel Biology Past Exam Papers (2B101)
June 2017 Biology/ Science Unit B1: Influences on Life (5BI1F/01) – Foundation Download Paper – Download Marking Scheme
June 2017 Biology/ Science Unit B1: Influences on Life (5BI1H/01) – Higher Download Paper – Download Marking Scheme
June 2017 Biology/ Additional Science Unit B2: The Components of Life (5BI2F/01) – Foundation Download Paper – Download Marking Scheme
June 2017 Biology/ Additional Science Unit B2: The Components of Life (5BI2H/01) – Higher Download Paper – Download Marking Scheme
June 2017 Biology Unit B3: Using Biology (5BI3F/01) – Foundation Download Paper – Download Marking Scheme
June 2017 Biology Unit B3: Using Biology (5BI3H/01) – Higher Download Paper – Download Marking Scheme
June 2016 Edexcel Biology Past Exam Papers (2B101)
June 2016 Biology/ Science Unit B1: Influences on Life (5BI1F/01) – Foundation Download Paper – Download Marking Scheme
June 2016 Biology/ Science Unit B1: Influences on Life (5BI1H/01) – Higher Download Paper – Download Marking Scheme
June 2016 Biology/ Additional Science Unit B2: The Components of Life (5BI2F/01) – Foundation Download Paper – Download Marking Scheme
June 2016 Biology/ Additional Science Unit B2: The Components of Life (5BI2H/01) – Higher Download Paper – Download Marking Scheme
June 2016 Biology Unit B3: Using Biology (5BI3F/01) – Foundation Download Paper – Download Marking Scheme
June 2016 Biology Unit B3: Using Biology (5BI3H/01) – Higher Download Paper – Download Marking Scheme
June 2015 (2BI01)
Biology/ Science Unit B1: Influences on Life (5BI1F/01) – Foundation – Download Paper – Download Marking Scheme
Biology/ Science Unit B1: Influences on Life (5BI1H/01) – Higher – Download Paper – Download Marking Scheme
Biology/ Additional Science Unit B2: The Components of Life (5BI2F/01) – Foundation Download Paper – Download Marking Scheme
Biology/ Additional Science Unit B2: The Components of Life (5BI2H/01) – Higher Download Paper – Download Marking Scheme
Biology Unit B3: Using Biology (5BI3F/01) – Foundation – Download Paper – Download Marking Scheme
Biology Unit B3: Using Biology (5BI3H/01) – Higher – Download Paper – Download Marking Scheme
June 2014 (2BI01)
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Biology library
Welcome to the biology library, unit 1: intro to biology, unit 2: chemistry of life, unit 3: water, acids, and bases, unit 4: properties of carbon, unit 5: macromolecules, unit 6: elements of life, unit 7: energy and enzymes, unit 8: structure of a cell, unit 9: more about cells, unit 10: membranes and transport, unit 11: more about membranes, unit 12: cellular respiration, unit 13: photosynthesis, unit 14: cell signaling, unit 15: cell division, unit 16: classical and molecular genetics, unit 17: dna as the genetic material, unit 18: central dogma (dna to rna to protein), unit 19: gene regulation, unit 20: biotechnology, unit 21: more molecular biology, unit 22: developmental biology, unit 23: bacteria and archaea, unit 24: viruses, unit 25: evolution and the tree of life, unit 26: more about evolution and natural selection, unit 27: history of life on earth, unit 28: ecology, unit 29: more about ecology, unit 30: biodiversity and conservation, unit 31: behavioral biology, unit 32: principles of physiology, unit 33: human biology, unit 34: plant biology, unit 35: ap free response worked examples, unit 36: crash course: biology and ecology, unit 37: meet the biology professional.
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Erin Dolan Receives Meigs Distinguished Teaching Professorship
Erin Dolan, professor in Biochemistry and Molecular Biology, has received a 2024 Meigs Distinguished Teaching Professorship for her excellence in teaching. Dolan was recognized for prioritizing students learning science by thinking scientifically, making scientific careers attainable via research mentoring, and scaling access to research opportunities via course-based research. The Meigs Professorship communicates the University of Georgia’s commitment to excellence in teaching, the value placed on the learning experiences of our students, and the centrality of instruction to the University's mission. It is named for Josiah Meigs, who presided over the University’s first class of graduates.
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The beauty of biology
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When Hanjun Lee arrived at MIT, he was set on becoming a Course 5 chemistry student. Based on his experience in high school, biology was all about rote memorization.
That changed when he took course 7.03 (Genetics) , taught by then-professor Aviv Regev , now head and executive vice president of research and early development at Genentech, and Peter Reddien , professor of biology and core member and associate director of the Whitehead Institute for Biomedical Research.
He notes that friends from other schools don’t cite a single course that changed their major, but he’s not alone in choosing Course 7 because of 7.03.
“Genetics has this interesting force, especially in MIT biology. The department’s historical — and active — role in genetics research ties directly into the way the course is taught,” Lee says. “Biology is about logic, scientific reasoning, and posing the right questions.”
A few years later, as a teaching assistant for class 7.002 (Fundamentals of Experimental Molecular Biology ), he came to value how much care MIT biology professors take in presenting the material for all offered courses.
“I really appreciate how much effort MIT professors put into their teaching,” Lee says. “As a TA, you realize the beauty of how the professors organize these things — because they’re teaching you in a specific way, and you can grasp the beauty of it — there’s a beauty in studying and finding the patterns in nature.”
An undertaking to apply
To attend MIT at all hadn’t exactly been a lifelong dream. In fact, it didn’t occur to Lee that he could or should apply until he represented South Korea at the 49th International Chemistry Olympiad, where he won a Gold Medal in 2017. There, he had the chance to speak with MIT alumni, as well as current and aspiring students. More than half of those aspiring students eventually enrolled, Lee among them.
“Before that, MIT was this nearly mythical institution, so that experience really changed my life,” Lee recalls. “I heard so many different stories from people with so many different backgrounds — all converging towards the same enthusiasm towards science.”
At the time, Lee was already attending medical school — a six-year undergraduate program in Korea — that would lead to a stable career in medicine. Attending MIT would involve both changing his career plans and uprooting his life, leaving all his friends and family behind.
His parents weren’t especially enthusiastic about his desire to study at MIT, so it was up to Lee to meet the application requirements. He woke up at 3 a.m. to find his own way to the only SAT testing site in South Korea — an undertaking he now recalls with a laugh. In just three months, he had gathered everything he needed; MIT was the only institution in the United States Lee applied to.
He arrived in Cambridge, Massachusetts, in 2018 but attended MIT only for a semester before returning to Korea for his two years of mandatory military service.
“During military service, my goal was to read as many papers as possible, because I wondered what topic of science I’m drawn to — and many of the papers I was reading were authored by people I recognized, people who taught biology at MIT,” Lee says. “I became really interested in cancer biology.”
Return to MIT
When he returned to campus, Lee pledged to do everything he could to meet with faculty and discuss their work. To that end, he joined the MIT Undergraduate Research Journal , allowing him to interview professors. He notes that most MIT faculty are enthusiastic about being contacted by undergraduate students.
Stateside, Lee also reached out to Michael Lawrence , an assistant professor of pathology at Harvard Medical School and assistant geneticist at Mass General Cancer Center, about a preprint concerning APOBEC, an enzyme Lee had studied at Seoul National University. Lawrence’s lab was looking into APOBEC and cancer evolution — and the idea that the enzyme might drive drug resistance to cancer treatment.
“Since he joined my lab, I’ve been absolutely amazed by his scientific talents,” Lawrence says. “Hanjun’s scientific maturity and achievements are extremely rare, especially in an undergraduate student.”
Lee has made new discoveries from genomic data and was involved in publishing a paper in Molecular Cell and a paper in Nature Genetics . In the latter, the lab identified the source of background noise in chromosome conformation capture experiments, a technique for analyzing chromatin in cells.
Lawrence thinks Lee “is destined for great leadership in science.” In the meantime, Lee has gained valuable insights into how much work these types of achievements require.
“Doing research has been rewarding, but it also taught me to appreciate that science is almost 100 percent about failures,” Lee says. “It is those failures that end up leading you to the path of success.”
Widening the scope
Lee’s personal motto is that to excel in a specific field, one must have a broad sense of what the entire field looks like, and suggests other budding scientists enroll in courses distant from their research area. He also says it was key to see his peers as collaborators rather than competitors, and that each student will excel in their own unique way.
“Your MIT experience is defined by interactions with others,” Lee says. “They will help identify and shape your path.”
For his accomplishments, Lee was recently named an American Association for Cancer Research Undergraduate Scholar . Last year, he also spoke at the Gordon Research Conference on Cell Growth and Proliferation about his work on the retinoblastoma gene product RB. Lee was also among the 2024 Biology Undergraduate Award Winners, recognized with the Salvador E. Luria Prize for outstanding scholarship and research of publication quality.
Encouraged by positive course evaluations during his time as a TA, Lee hopes to inspire other students in the future through teaching. Lee has recently decided to pursue a PhD in cancer biology at Harvard Medical School, although his interests remain broad.
“I want to explore other fields of biology as well,” he says. “I have so many questions that I want to answer.”
Although initially resistant, Lee’s mother and father are now “immensely proud to be MIT parents” and will be coming to Cambridge in May to celebrate Lee’s graduation.
“Throughout my years here, they’ve been able to see how I’ve changed,” he says. “I don’t think I’m a great scientist, yet, but I now have some sense of how to become one.”
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Revision Notes. Concise resources for the Edexcel GCSE Biology course. 1. Key Concepts in Biology. 1.1 Cell Structure. 1.1.1 Eukaryotic Organisms. 1.1.2 Eukaryotic Organisms: Animals & Plants. 1.1.3 Eukaryotic Organisms: Fungi & Protoctists. 1.1.4 Prokaryotic Organisms.
This interactive quiz is suitable for GCSE Biology (single science) students studying communicable diseases and how the body and medicine work to prevent them. Plant disease - Edexcel. Pathogens ...
FREE Biology revision notes on Eukaryotic Organisms. Designed by the teachers at SAVE MY EXAMS for the Edexcel GCSE Biology syllabus.
Content and assessment overview. The Pearson Edexcel Level 1/Level 2 GCSE (9- 1) in Biology consists of two externally-examined papers. These are available at foundation tier and higher tier. Students must complete all assessments in the same tier. Students must complete all assessment in May/June in any single year.
A playlist that covers the entire of the Edexcel IGCSE Biology 9-1 course!
These papers are essential resources for students preparing for their GCSE Biology exams. Practicing with these past papers can help improve your exam technique, enhance your knowledge and understanding of the subject matter, and boost your confidence. Our page provides access to a range of past papers, including those from the official Edexcel ...
Assessment. These resources have been written to support the Pearson Edexcel International GCSE (9-1) specification, a linear qualification which consists of examinations at the end of the course of study for Biology, Chemistry, Physics, Single Award Science and Double Award Science.
Download Paper - Download Marking Scheme. November 2020 Biology Paper 2 (1BI0/2H) - Higher Tier. Download Paper - Download Marking Scheme. Some of the above papers are labelled June 2020. June 2019 Pearson Edexcel Biology Past Exam Papers (1BI0) June 2019 Biology Paper 1 (1BI0/1F) - Foundation Tier. Download Paper - Download Marking ...
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Use the formula for the area of a circle (πr2) to calculate the area of these circles. 5. Divide the area of the smaller circle by the larger, and multiply by 100. This is the percentage of the bacterial culture that has been destroyed by the antibiotic. The higher the percentage, the more effective the antibiotic.
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Cycles within Ecosystems. Human Influences on the Environment. 5. Use of Biological Resources. Food Production. Selective Breeding. Genetic Modification (Genetic Engineering) Cloning. Revision notes for the Edexcel IGCSE Biology syllabus, written by the Biology experts at Save My Exams.
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Erin Dolan, professor in Biochemistry and Molecular Biology, has received a 2024 Meigs Distinguished Teaching Professorship for her excellence in teaching. Dolan was recognized for prioritizing students learning science by thinking scientifically, making scientific careers attainable via research mentoring, and scaling access to research opportunities via course-based research.
The beauty of biology. Senior Hanjun Lee planned to pursue chemistry at MIT. A course in genetics changed that. Hanjun Lee will pursue a PhD in cancer biology at Harvard Medical School this fall. Photo courtesy of Hanjun Lee. An illustration for the paper "Chromatin-bound RB targets promoters, enhancers, and CTCF-bound loci and is ...